When the Besties decide to track their new and incredibly mean sub-teacher, Ms. Lemon, to her lemon-shaped house, they discover that she is no ordinary human being. They also find out that she not only hates children, but also trees and is determined to destroy the ideal and environmentally friendly village of Rosemallow. As the Besties go through funny and unbelievable adventures, they will try their best to stop Ms. Lemon’s wicked plan